How to combine LTO with symbol versioning

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Hello,

I would like to compile a shared library using both symbol versioning
and link-time optimization (LTO). However, as soon as I turn on LTO,
some of the exported symbols vanish. For example:

Start by defining two implementations of a function fun:

$ cat fun.c 
#include <stdio.h>

int fun1(void);
int fun2(void);

__asm__(".symver fun1,fun@v1");
int fun1() {
    printf("fun1 called\n");
    return 1;
}

__asm__(".symver fun2,fun@@v2");
int fun2() {
    printf("fun2 called\n");
    return 2;
}

Create a version script to ensure that only fun is exported:

$ cat versionscript 
v1 {
    global:
        fun;
    local:
        *;
};
v2 {
    global:
        fun;
} v1;

First attempt, compile without LTO:

$ gcc -o fun.o -Wall -Wextra -O2 -fPIC -c fun.c
$ gcc -o libfun.so.1 -shared -fPIC -Wl,--version-script,versionscript fun.o
$ nm -D --with-symbol-versions libfun.so.1 | grep fun
00000000000006b0 T fun@@v2
0000000000000690 T fun@v1

..exactly as it should be. But if I compile with LTO:

$ gcc -o fun.o -Wall -Wextra -flto -O2 -fPIC -c fun.c
$ gcc -o libfun.so.1 -flto -shared -fPIC -Wl,--version-script,versionscript fun.o
$ nm -D --with-symbol-versions libfun.so.1 | grep fun

..no symbols exported anymore.

What am I doing wrong?

$ gcc --version
gcc (Debian 6.3.0-18) 6.3.0 20170516

Best,
-Nikolaus
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